Associate Professor Chao, Yi-Shan

PhD, Department of Biological Sciences, National Sun Yat-sen University, R.O.C.
Courses offered:
Evolutionary biology, Genetic diversity, General Biology, Plant ecology, Plant morphology and systematics
Aquatic plants, Carnivorous plants, Plant taxonomy, Plant systematics and evolution, Pteridophytes


Journal Publications

  1. Chao, Yi-Shan*. 2019. New record of Pteris esquirolii in Taiwan, with two similar new species, Pteris rugosifolia and Pteris subesquirolii. Systematic Botany 44 (2): in press. (2017 IF 1.515; rank 100/223 plant sciences)

  2. Chao, Yi-Shan*Yao-Moan Huang. 2018. Spore morphology and its systematic implication in Pteris (Pteridaceae). PLOS ONE13(11): e0207712. (2017 IF 2.766rank 15/64 multidisciplinary sciences)

  3. Dong, Shi-Yong, Cheng-Wei Chen, Shi-Shi Tan, Hui-Guo Zhao, Zheng-Yu Zuo, Yi-Shan Chao, and Yi-Han Chang. 2018. New insights on the phylogeny of Tectaria (Tectariaceae), with special reference to Polydictyum as a distinct lineage. Journal of Systematics and Evolution 56 (2): 139-147. (2017 IF 3.657; rank 26/223 plant sciences)

  4. Chao, Yi-ShanAndi Maryani A. Mustapeng, Cheng-Wei Chen, Wen-Liang Chiou*. 2017. Pteris borneensis (Pteridaceae), a new species from Borneo, with re-circumscription of Pteris decrescens and Pteris parviloba. Systematic Botany 42(4):1-9. (2016 IF 1.240; rank 116/212 plant sciences)

  5. Chao, Yi-ShanAtsushi Ebihara, Wen-Liang Chiou, Yao-Moan Huang*. 2017. Pteris latipinna sp. nov. (Pteridaceae), a new species segregated from Pteris fauriei. Phytokeys 85: 95–108. (2016 IF 1.116; rank 127/212 plant sciences)

  6. Fujiwara, Tao, Ayumi Uehara, Tsukasa Iwashina, Sadamu Matsumoto, Yi-Han Chang, Yi-Shan Chao, and Yausyuki Watano*. 2017. Allotetraploid cryptic species in Asplenium normale in the Japanese Archipelago, detected by chemotaxonomic and multilocus genotype approaches. American Journal of Botany104(9): 1 – 17(2016 IF 3.050; rank 39/212 plant sciences)

  7. Chen, Cheng-Wei, Stuart Lindsay, Li-Yaung Kuo, Christopher R. Fraser-Jenkins, Atsushi Ebihara, Hong Truong Luu, Chong Wook Park, Yi-Shan Chao*, Yao-Moan Huang*, and Wen-Liang Chiou. 2017. A systematic study of East Asian vittarioid ferns (Pteridaceae: Vittarioideae). Botanical Journal of the Linnean Society 183:545-560. (2016 IF 2.277; rank 62/212 plant sciences)

  8. Hsu, Tinan-Chuan, Zhi-Hao ChenYi-Shan Chao*. 2017.New additions of the bladderworts (Lentibulariaceae) in Taiwan. Taiwania 62 (1): 99104. (2017前無資料;2017 IF 0.421rank 29/37 Horticulture)

  9. The Pteridophyte Phylogeny Group (including Yi-Shan Chao). 2016. A community-derived classification for extant lycophytes and ferns. Journal of Systematics and Evolution 54(6): 563-603. (2015 IF 1.134; rank 123/209 plant sciences)

  10. Dong, Shi-Yong, Zheng-Yu Zuo, Yi-Shan Chao, Kipiro Damas, Bernard Sule. 2016. New species of the fern genus Lindsaea (Lindsaeaceae) from New Guinea with notes on the phylogeny of L. sect. Synaphlebium. PLOS ONE 11(10): e0163686. (2015 IF 3.057rank 11/63 multidisciplinary sciences)

  11. Chao, Yi-Shan, Ho-Yih Liu, and Wen-Liang Chiou*. 2015. Taxonomic revision of the Pteris cadieri complex (Pteridaceae).Phytotaxa 230: 130-150. (2014 IF 1.318; rank 105/204 plant sciences )

  12. Ding, Hui-Hui (equal contributor), Chao, Yi-Shan (equal contributor), Callado, John Rey, Dong, Shi-Yong, 2014. Phylogeny and character evolution of the fern genus Tectaria (Tectariaceae) in the Old World inferred from chloroplast DNA sequences. Molecular Phylogenetics and Evolution 80: 66-78.
  13. Chao, Yi-Shan, Germinal Rouhan, Victor B. Amoroso, and Wen-Liang Chiou*. 2014. Molecular phylogeny and biogeography of the fern genus Pteris (Pteridaceae). Annals of Botany 114: 109–124.
  14. Kao, Tzu-Tong, Wen-Liang Chiou, Yuan-Hsu Sheng, Chun-Ming Chen, Yi-Shan Chao, Yao-Moan Huang*. 2014. Hybrid Origins of Nephrolepis ×hippocrepicis Miyam. (Nephrolepidaceae). The International Journal of Plant Reproductive Biology 6: 1-14.
  15. Ebihara, Atsushi*, Narumi Nakato, Sadamu Matsumoto, Yi-Shan Chao, and Li-Yaung Kuo. 2014. Cytotaxonomic Studies on Thirteen Ferns of Taiwan. Bulletin of the National Science Museum. Series B 40: 19-28. 
  16. Chao, Yi-Shan, Atsushi Ebihara, Yi-Han Chang, Ri-Hong Jiang, Lu Thi Ngan, and Wen-Liang Chiou*. 2013. New distribution of Pteris kawabatae Sa. Kurata and re-circumscription of Pteris arisanensis Tagawa. Taiwania 58: 305-310.
  17. Ding, Hui-Hui, Yi-Shan Chao, and Shi-Yong Dong*. 2013 Taxonomic novelties in the fern genus Tectaria (Tectariaceae). Phytotaxa, 122: 61-64.
  18. Chao, Yi-ShanYa-Nan Wang, Chieh-Ting Wang*. 2012. A DNA barcode for forest. Journal of the Experimental Forest of National Taiwan University 26: 299-304.
  19. Chao, Yi-Shan, Shi-Yong Dong, Yu-Chung Chiang, Ho-Yih Liu, and Wen-Liang Chiou*. 2012. Extreme multiple reticulate origins of the Pteris cadieri complex. International Journal of Molecular Sciences 13: 4523-4544. 
  20. Chao, Yi-Shan, Yi-Han Chang, Lu Thi Nga, Do Van Truong, Ho-Yih Liu, and Wen-Liang Chiou*. 2012. Different ploids of Pteris grevilleana Wall. ex Agardh var. ornata Alderw. (Pteridaceae) in Taiwan and Vietnam. Taiwania, 57:278-282.
  21. Chao, Yi-Shan, Ho-Yih Liu, Yu-Chung Chiang, and Wen-Liang Chiou*. 2012 Polyploidy and speciation in the Pteris (Pteridaceae). Journal of Botany, vol. 2012, Article ID 817920, 7 pages. 
  22. Dong, Shi-Yong*, R. Mujahidin, Ling-Ling Wei, Yi-Shan Chao. 2012. A new species of Asplenium sect. Thamnopteris (Aspleniaceae) from Indonesia. Blumea 57: 190-194. 
  23. Ju, Li-Ping (equal contributor), Chia-Chi Kuo, Yi-Shan Chao (equal contributor), Yu-Pin Cheng, Xun Gong, and Yu-Chung Chiang*. 2011. Microsatellite primers in the native perennial cycad, Cycas taitungensis (Cycadaceae). American Journal of Botany, e84-e86. 
  24. Chao, Yi-Shan, Ho-Yih Liu, Yao-Moan Huang, and Wen-Liang Chiou*. 2010. Reproductive traits of Pteris cadieri and P. grevilleana in Taiwan: Implications for their hybrid origin. Botanical Studies, 51: 209-216. 
  25. Yang, Yuen-Po, Ho-Yih Liu, and Yi-Shan Chao* (corresponding author). 2009. Trap gland morphology and its systematic implications in Taiwan Utricularia (Lentibulariaceae). Flora, 204: 692-699. 
  26. 張惠珠、趙怡姍、柯智仁、楊遠波*。2006。墾丁國家公園水生植物物種調查。國家公園學報 16: 19-31。

Other publication

  1. Liao, Pei-Chun, Yi-Shan Chao, and Yu-Chung Chiang. 2012. The species occurrence, hybridization and speciation and in postglacial East and South Asia. In: The Molecular Basis of Plant Genetic Diversity, Mahmut Çalışkan ed. pp. 33-60. InTech Open Access Publisher, Hatay, Turkey.
  2. 楊遠波、趙怡姍。2006。墾丁國家公園水生植物圖鑑。內政部營建署墾丁國家公園管理處。恆春。

Symposium publication:

  1. 丁惠惠、趙怡姍、董仕勇。2012。東亞叉蕨屬的分子系統學研究。中國蕨類學術研討會。中國,海口。
  2. Chao, Yi-Shan, and Yu-Chung Chiang, 2011. Molecular evolution and genetic diversity of Calmodulin gene family in Oryza L. 18th International Botanical congress, Melbourne, Australia.
  3. Chao, Yi-Shan, Shi-Yong Dong, Yu-Chung Chiang, Ho-Yih Liu, and Wen-Liang Chiou. 2011. Apogamous ferns with extreme morphological variation: reticulate evolution of Pteris cadieri complex. 18th International Botanical congress, Melbourne, Australia.
  4. Chao, Yi-Shan, Shi-Yong Dong, Yu-Chung Chiang, Ho-Yih Liu, and Wen-Liang Chiou. 2010. Insight into origins of the Pteris cadieri complex. 5th Symposium on Asian Pteridology, Guangzhou, China.
  5. Chao, Yi-Shan, Yu-Chung Chiang, Yao-Moan Huang, Shi-Yong Dong, Ho-Yih Liu, and Wen-Liang Chiou. 2010. Problematic taxonomy and phylogeny in genus Pteris (Pteridaceae). 8th International Flora Malesiana Symposium. Singapore Botanical Garden. Singapore.
  6. Liao, Pei-Chun, , Yi-Shan Chao, Yu-Chung Chiang. 2010. Multiple case studies of phytogeography in postglacial east and south Asia. International Symposium on Forest Biodiversity & Conservation. Nantou, Taiwan. 14-28.
  7. 趙怡姍、劉和義、邱文良。2010。蕨類的網狀演化:以二形鳳尾蕨複合群為例。2010植物多樣性與系統分類研討會。台灣植物分類學會。台北
  8. 趙怡姍、劉和義、邱文良。2009。二形鳳尾蕨複合種群是由多次雜交形成的雜交群。2009 植物多樣性與系統分類研討會。台灣植物分類學會。台北
  9. 趙怡姍、劉和義、邱文良。2008。二形鳳尾蕨蕨複合群到底有多少分類群?葉綠體DNA資料分析。2008 植物多樣性與系統分類研討會。台灣植物分類學會。台北。
  10. Chao, Yi-Shan, Ho-Yih Liu, and Wen-Liang Chiou. 2007. The value of C-Value in Pteris cadieri complex. 4th Symposium on Asian Pteridology. Central Mundanao University, Musuan, Bukidnon, Philippines.
  11. 趙怡姍、劉和義、邱文良。2007。C-value在二形鳳尾蕨分類研究之應用。植物多樣性與系統分類研討會。台灣植物分類學會。台北。
  12. Chao, Yi-Shan, Ho-Yih Liu and Yuen-Po Yang. 2006. Trap gland morphology and its systematic implication in Taiwan Utricularia. The 6th Conference of International Carnivorous Plant Society. Maryland, USA.
  13. 趙怡姍、林素貞。2000。台灣地區橡膠製品業能源消費與二氧化碳減量分析。第十三屆環境規劃與管理研討會。
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